2-tissue-2.ppt
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1、筛管筛管sieve tube sieve tube 伴胞伴胞companion cellcompanion cell筛管筛管被子植物的韧皮部被子植物的韧皮部,运输运输有机物有机物.筛管筛管由管状的由管状的无细胞无细胞核的活细胞核的活细胞连接成连接成组成筛管的每个细胞组成筛管的每个细胞叫做叫做筛管分子筛管分子 sieve elements1筛管分子的细胞壁为筛管分子的细胞壁为初生壁初生壁性质。性质。端壁称为端壁称为筛板筛板;分布有小孔分布有小孔-筛孔筛孔。2筛板和筛孔筛板和筛孔Mature sieve-tube members from thephloem in the stem of corn
2、 and squash.34P-proteinCallose筛管运送养分的速度筛管运送养分的速度10-100cm/h56受伤时,通过筛孔的原生质形成受伤时,通过筛孔的原生质形成联络索联络索阻断相邻的筛管分子阻断相邻的筛管分子筛孔纵切面筛孔纵切面(侧面观)(侧面观)7筛管分子的老化胼胝质沉积在筛板上,筛管分子的老化胼胝质沉积在筛板上,形成形成胼胝体胼胝体阻断筛孔,失去输导功能。阻断筛孔,失去输导功能。8胼胝体(绿色荧光)胼胝体(绿色荧光)9P-protein(phloem protein)A protein found in large amounts in the sap-conducting
3、 sieve elements of phloem tissue in plants.It takes various forms in the mature sieve element,depending on plant species,ranging from a network of filaments to discrete crystalline bodies.One general property is its ability to form a gel,and it functions as a puncture repair substance,forming a plug
4、 at any site of damage in the sieve element,thus preventing loss of food materials being translocated by the phloem.In an intact functioning sieve element,the P-protein is mainly found lining the interior wall.10Longitudinal section of sieve tube members in stem of cucumber(Cucumis).This material wa
5、s fixed in such a way as to enhance the formation of P-protein plugs,two of which are present here,stained red.11筛管分子侧壁上存在着初生纹孔场筛管分子侧壁上存在着初生纹孔场筛域筛域 12伴胞伴胞-筛管旁边的一个或多个小细胞。筛管旁边的一个或多个小细胞。The phloem in the stem of squash南瓜南瓜13sieve tube members can have multiple companion cells 14筛管分子与伴胞之间筛管分子与伴胞之间具有筛域和胞
6、间连丝具有筛域和胞间连丝?15筛管和伴胞由一个细胞不等分裂产生筛管和伴胞由一个细胞不等分裂产生 16早早期期筛筛管管分分子子伴伴胞胞171819Sieve tube members and companion cells typically function for only a few weeks or months,then are replaced by new cells.20Transverse section of phloem in a cactus(Espostoopsis dybowskii).the phloem of this species is trapped bet
7、ween the expanding xylem21Transverse section of old stem of grape(Vitis)from a stem that was much older;the sieve tube members and companion cells have collapsed into the solid,dark red mass 22筛胞筛胞 蕨类和裸子植物体内输导有机物的细胞蕨类和裸子植物体内输导有机物的细胞。Transverse section of a fern rhizome(Pteridium aquilinum).The cente
8、r of this micrograph is occupied by xylem(the large red-stained cells),and surrounding that is a band of large,empty-looking cells with thin walls(arrows);those are the sieve cells of this fern(蕨类蕨类)phloem.23筛管筛管被子植物韧皮部被子植物韧皮部呈管道状,管径较大呈管道状,管径较大端壁有筛板端壁有筛板,有伴胞有伴胞输导能力较强输导能力较强筛胞筛胞蕨类,裸子植物韧皮部蕨类,裸子植物韧皮部呈细胞
9、状,管径小,呈细胞状,管径小,尖削端相接无筛板无伴胞尖削端相接无筛板无伴胞输导功能较弱输导功能较弱24(三)机械组织(三)机械组织 mechanical tissuemechanical tissue细胞壁细胞壁均匀或不均匀次生均匀或不均匀次生加厚。加厚。25机械组织根据形态及细胞机械组织根据形态及细胞壁的加厚方式,可分为壁的加厚方式,可分为厚角组织厚角组织 collenchymacollenchyma厚壁组织厚壁组织 scleranchymascleranchyma261 1厚角组织厚角组织 活细胞活细胞 CollenchymaCollenchyma 角隅处增厚角隅处增厚,是初生壁性质是初生
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